Technical papers 1999-present, with pdfs (or links to pdfs) where available

Naish, D. & Perron, R. 2014. Structure and function of the cassowary’s casque and its implications for cassowary history, biology and evolution. Historical Biology 10.1080/08912963.2014.985669 [pdf here]

Young, M. T., Steel, L., Foffa, D., Price, T., Naish, D. & Tennant, J. P. 2014. Marine tethysuchian crocodyliform from the ?Aptian-Albian (Lower Cretaceous) of the Isle of Wight, UK. Biological Journal of the Linnean Society 113, 854-871. [pdf here]

Lee, M. S. Y., Cau, A., Naish, D. & Dyke, G. J. 2014. Sustained miniaturization and anatomical innovation in the dinosaurian ancestors of birds. Science 345, 562-565. [pdf here]

Fischer, V., Arkangelsky, M. S., Naish, D., Stenshin, I. M., Uspensky, G. N. & & Godefroit, P. 2014. Simbirskiasaurus and Pervushovisaurus reassessed: implications for the taxonomy and cranial osteology of Cretaceous platypterygiine ichthyosaurs. Zoological Journal of the Linnean Society 171, 822-841. [pdf here]

Lee, M. S. Y., Cau, A., Naish, D. & Dyke, G. 2014. Morphological clocks in paleontology, and a Mid-Cretaceous origin of crown Aves. Systematic Biology 63, 442-449. [pdf here]

Naish, D., Sakamoto, M., Hocking, P. & Sanchez, G. 2014. ‘Mystery big cats’ in the Peruvian Amazon: morphometrics solve a cryptozoological mystery. PeerJ 2:e291; DOI 10.7717/peerj.291

Vremir, M., Bălc, R., Csiki-Sava, Z. Brusatte, S. L., Dyke, G., Naish, D., Norell, M. A. 2014. Petreşti-Arini – An important but ephemeral Upper Cretaceous continental vertebrate site in the southwestern Transylvanian Basin, Romania. Cretaceous Research 49, 13-38. [pdf here]

Naish, D. 2014. The fossil record of bird behaviour. Journal of Zoology 292, 268-280. [pdf here]

Brusatte, S. L., Vremir, M., Watanabe, A., Csiki-Sava, Z., Naish, D., Dyke, G., Erickson, G. M. & Norell, M. A. 2013. An infant ornithopod dinosaur tibia from the Late Cretaceous of Sebeş, Romania. Terra Sebus. Acta Musei Sabesiensis 5, 627-644. [pdf here]

Witton, M. P. & Naish, D. 2013. Azhdarchid pterosaurs: water-trawling pelican mimics or “terrestrial stalkers”? Acta Palaeontologica Polonica doi:

Dyke, G., de Kat, R., Palmer, C., van der Kindere, J., Naish, D. & Ganapathisubramani, B. 2013. Aerodynamic performance of the feathered dinosaur Microraptor and the evolution of feathered flight. Nature Communications 4, Article number: 2489 doi:10.1038/ncomms3489 [pdf here]

Fischer, V., Appleby, R. M., Naish, D., Liston, J., Riding, J. B., Brindley, S. & Godefroit, P. 2013. A basal thunnosaurian from Iraq reveals disparate phylogenetic origins for Cretaceous ichthyosaurs. Biology Letters 9, 20130021 [pdf here]

Blake, M., Naish, D., Larson, G., King, C. L., Nowell, G., Sakamoto, M. & Barnett, R. 2013. Multidisciplinary investigation of a ‘British big cat’: a lynx killed in southern England c. 1903. Historical Biology doi:10.1080/08912963.2013.785541 [pdf here]

Hone, D. W. E. & Naish, D. 2013. The ‘species recognition hypothesis’ does not explain the presence and evolution of exaggerated structures in non-avialan dinosaurs. Journal of Zoology doi:10.1111/jzo.12035 [pdf here]

Knell, R. J., Naish, D., Tomkins, J. L. & Hone, D. W. E. 2013. Is sexual selection defined by dimorphism alone? A reply to Padian and Horner. Trends in Ecology and Evolution xx. [pdf here]

Naish, D., Simpson, M. I. & Dyke, G. J. 2013. A new small-bodied azhdarchoid pterosaur from the Lower Cretaceous of England and its implications for pterosaur anatomy, diversity and phylogeny. PLoS ONE 8(3): e58451. doi:10.1371/journal.pone.0058451

Vremir, M., Kellner, A. W. A., Naish. D. & Dyke, G. J. 2013. A new azhdarchid pterosaur from the Late Cretaceous of the Transylvanian Basin, Romania: implications for azhdarchid diversity and distribution. PLoS ONE 8(1): e54268. doi:10.1371/journal.pone.0054268

Benson, R. B. J., Ketchum, H. F., Naish, D. & Turner, L. E. 2012. A new leptocleidid (Sauropterygia, Plesiosauria) from the Vectis Formation (Early Barremian-early Aptian; Early Cretaceous) of the Isle of Wight and the evolution of Leptocleididae, a controversial clade. Journal of Systematic Palaeontology DOI: 10.1080/14772019.2011.634444 [pdf here]

Birn-Jeffery, A. V., Miller, C. E., Naish, D., Rayfield, E. J., Hone, D. W. E. 2012. Pedal claw curvature in birds, lizards and Mesozoic dinosaurs – complicated categories and compensating for mass-specific and phylogenetic control. PLoS ONE 7(12): e50555. doi:10.1371/journal.pone.0050555

Naish, D. 2012. Identifying ‘Jaws’, the Margaret River mammal carcase. The Journal of Cryptozoology 1, 45-55.

Woodley, M. A., McCormick, C. A. & Naish, D. 2012. Response to Bousfield & LeBlond: Shooting pipefish in a barrel; or sauropterygian “mega-serpents” and Occam’s razor. Journal of Scientific Exploration 26, 151-154.

Knell, R. J., Naish, D., Tomkins, J. L. & Hone, D. W. E. 2012. Sexual selection in prehistoric animals: detection and implications. Trends in Ecology and Evolution 28, 38-47. [pdf here]

Dyke, G., Vremir, M., Kaiser, G. & Naish, D. 2012. A drowned Mesozoic bird breeding colony from the Late Cretaceous of Transylvania. Naturwissenschaften 99, 435-442. [pdf here]

Fischer, V., Maisch, M. W., Naish, D., Kosma, R., Liston, J., Joger, U., Krüger, F. J., Pérez, J. P., Tainsh, J. & Appleby, R. M. 2012. New ophthalmosaurid ichthyosaurs from the European Lower Cretaceous demonstrate extensive ichthyosaur survival across the Jurassic-Cretaceous boundary. PLoS ONE 7(1): e29234. doi:10.1371/journal.pone.0029234

Dyke, G. J. & Naish, D. 2011. What about European alvarezsauroids? Proceedings of the National Academy of Sciences of the United States of America doi:10.1073/pnas.1101602108 [pdf here]

Naish, D. & Sweetman, S. C. 2011. A tiny maniraptoran dinosaur in the Lower Cretaceous Hastings Group: evidence from a new vertebrate-bearing locality in south-east England. Cretaceous Research 32, 464-471. [pdf here]

Taylor, M. P., Hone, D. W. E., Wedel, M. J., & Naish, D. 2011. The long necks of sauropods did not evolve primarily through sexual selection. Journal of Zoology 285, 150-161. [pdf here]

Woodley, M. A., Naish, D., & McCormick, C. A. 2011. A baby sea-serpent no more: reinterpreting Hagelund’s juvenile “cadborosaur” report. Journal of Scientific Exploration 25, 495-512. [pdf here]

Hone, D. W. E., Naish, D. & Cuthill, I. C. 2011. Does mutual sexual selection explain the evolution of head crests in pterosaurs and dinosaurs? Lethaia 45, 139-156. [pdf here]

Naish, D., Dyke, G., Cau, A., Escuillié, F. & Godefroit, P. 2012. A gigantic bird from the Upper Cretaceous of Central Asia. Biology Letters 8, 97-100. [pdf here]

Dyke, G., Benton, M., Posmosanu, E., & Naish, D. 2010. Early Cretaceous (Berriasian) birds and pterosaurs from the Cornet bauxite mine, Romania. Palaeontology 54, 79-95. [pdf here]

Henderson, D. M. & Naish, D. 2010. Predicting the buoyancy, equilibrium and potential swimming ability of giraffes by computational analysis. Journal of Theoretical Biology 265, 151-159. [pdf here]

Taylor, M. P., Wedel, M. J. & Naish, D. 2009. Head and neck posture in sauropod dinosaurs inferred from extant animals. Acta Palaeontologica Polonica 54, 213-220. [pdf here]

Benson, R. B. J., Brusatte, S. L., Hutt, S. & Naish, D. 2009. A new large basal tetanuran (Dinosauria: Theropoda) from the Wessex Formation (Barremian) of the Isle of Wight, England. Journal of Vertebrate Paleontology 29, 612-615. [pdf here]

Witton, M. P. & Naish, D. 2008. A reappraisal of azhdarchid pterosaur functional morphology and paleoecology. PLoS ONE 3 (5): e2271. doi:10.1371/journal.pone.0002271

Woodley, M. A., Naish, D. & Shanahan, H. P. 2008. How many extant pinniped species remain to be described? Historical Biology 20, 225-235. [pdf here]

Naish, D. & Martill, D. M. 2008. Dinosaurs of Great Britain and the role of the Geological Society of London in their discovery: Ornithischia. Journal of the Geological Society, London 165, 613-623. [pdf here]

Sánchez-Hernández, B., Benton, M. J. & Naish, D. 2007. Dinosaurs and other fossil vertebrates from the Late Jurassic and Early Cretaceous of the Galve area, NE Spain. Palaeogeography, Palaeoclimatology, Palaeoecology 249, 180-215. [pdf here]

Taylor, M. P. & Naish, D. 2007. An unusual new neosauropod dinosaur from the Lower Cretaceous Hastings Beds Group of East Sussex, England. Palaeontology 50, 1547-1564. [pdf here]

Naish, D. & Martill, D. M. 2007. Dinosaurs of Great Britain and the role of the Geological Society of London in their discovery: basal Dinosauria and Saurischia. Journal of the Geological Society, London 164, 493-510. [pdf here]

Martill, D. M., Earland, S. & Naish, D. 2006. Dinosaurs in marine strata: evidence from the British Jurassic, including a review of the allochthonous vertebrate assemblage from the marine Kimmeridge Clay Formation (Upper Jurassic) of Great Britain. In Colectivo Arqueológico-Paleontológico Salense (ed) Actas de las III Jornadas sobre Dinosaurios y su Entorno. Salas de los Infantes (Burgos, España), pp. 47-83. [pdf here]

Martill, D. M. & Naish, D. 2006. Cranial crest development in the azhdarchoid pterosaur Tupuxuara, with a review of the genus and tapejarid monophyly. Palaeontology 49, 925-941. [pdf here]

Fielding, S., Martill, D. M. & Naish, D. 2005. Solnhofen-style soft-tissue preservation in a new species of turtle from the Crato Formation (Early Cretaceous, Aptian) of north-east Brazil. Palaeontology 48, 1301-1310. [pdf here]

Taylor, M. P. & Naish, D. 2005. The phylogenetic taxonomy of Diplodocoidea (Dinosauria: Sauropoda). PaleoBios 25, 1-7. [pdf here]

Naish, D., Martill, D. M. & Frey, E. 2004. Ecology, systematics and biogeographical relationships of dinosaurs, including a new theropod, from the Santana Formation (?Albian, Early Cretaceous) of Brazil. Historical Biology 16, 57-70. [pdf here]

Naish, D., Martill, D. M., Cooper, D. & Stevens, K. A. 2004. Europe’s largest dinosaur? A giant brachiosaurid cervical vertebra from the Wessex Formation (Early Cretaceous) of southern England. Cretaceous Research 25, 787-795. [pdf here]

Naish, D. 2004. New Zealand’s giant gecko: a review of current knowledge of Hoplodactylus delcourti and the kawekaweau of legend. The Cryptozoology Review 4 (2), 17-21.

Naish, D. & Dyke, G. J. 2004. Heptasteornis was no ornithomimid, troodontid, dromaeosaurid or owl: the first alvarezsaurid (Dinosauria: Theropoda) from Europe. Neues Jahrbuch für Geologie und Paläontologie, Monatshefte 2004, 385-401. [pdf here]

Naish, D. & Martill, D. M. 2003. Pterosaurs – a successful invasion of prehistoric skies. Biologist 50 (5), 213-216. [pdf here]

Naish, D. 2003. A definitive allosauroid (Dinosauria; Theropoda) from the Lower Cretaceous of East Sussex. Proceedings of the Geologists’ Association 114, 319-326. [pdf here]

Naish, D. 2002. The historical taxonomy of the Lower Cretaceous theropods (Dinosauria) Calamospondylus and Aristosuchus from the Isle of Wight. Proceedings of the Geologists’ Association 113, 153-163. [pdf here]

Walsh, S. A. & Naish, D. 2002. Fossil seals from late Neogene deposits in South America: a new pinniped (Carnivora, Mammalia) assemblage from Chile. Palaeontology 45, 821-842. [pdf here]

Naish, D. & Martill, D. M. 2002. A reappraisal of Thecocoelurus daviesi (Dinosauria: Theropoda) from the Early Cretaceous of the Isle of Wight. Proceedings of the Geologists’ Association 113, 23-30. [pdf here]

Hutt, S., Naish, D., Martill, D. M., Barker, M. J. & Newbery, P. 2001. A preliminary account of a new tyrannosauroid theropod from the Wessex Formation (Early Cretaceous) of southern England. Cretaceous Research 22, 227-242. [pdf here]

Naish, D. 2000. A small, unusual theropod (Dinosauria) femur from the Wealden Group (Lower Cretaceous) of the Isle of Wight, England. Neues Jahrbuch für Geologie und Paläontologie, Monatshefte 2000, 217-234. [pdf here]

Naish, D. 2000. Theropod dinosaurs in the trees: a historical review of arboreal habits amongst nonavian theropods. Archaeopteryx 18, 35-41. [pdf here]

Naish, D. 1999. Theropod dinosaur diversity and palaeobiology in the Wealden Group (Early Cretaceous) of England: evidence from a previously undescribed tibia. Geologie en Mijnbouw 78, 367-373. [pdf here]

Book chapters

Naish, D. 2013. South American native mammals. In McDade, M. (ed) Grzimek’s Animal Life Encyclopedia: Extinct Life. Gale Group (Farmington Mills, Michigan), pp. 567-576.

Naish, D. 2012. Birds. In Brett-Surman, M. K., Holtz, T. R. & Farlow, J. O. (eds) The Complete Dinosaur (Second Edition). Indiana University Press (Bloomington & Indianapolis), pp. 379-423.

Naish, D. 2011. Theropod dinosaurs. In Batten, D. J. (ed.) English Wealden Fossils. The Palaeontological Association (London), pp. 526-559.

Salisbury, S. W. & Naish, D. 2011. Crocodilians. In Batten, D. J. (ed.) English Wealden Fossils. The Palaeontological Association (London), pp. 305-369. [pdf here]

Naish, D. 2010. Pneumaticity, the early years: Wealden Supergroup dinosaurs and the hypothesis of saurischian pneumaticity. In Moody, R. T. J., Buffetaut, E., Naish, D. & Martill, D. M. (eds) Dinosaurs and Other Extinct Saurians: A Historical Perspective. Geological Society, London, Special Publications 343, pp. 229-236. [pdf here]

Moody, R. T. J. & Naish, D. 2010. Alan Jack Charig (1927-1997): an overview of his academic accomplishments and role in the world of fossil reptile research. In Moody, R. T. J., Buffetaut, E., Naish, D. & Martill, D. M. (eds) Dinosaurs and Other Extinct Saurians: A Historical Perspective. Geological Society, London, Special Publications 343, pp. 89-109. [pdf here]

Naish, D., Martill, D. M. & Merrick, I. 2007. Birds of the Crato Formation. In Martill, D. M., Bechly, G. & Loveridge, R. F. (eds) The Crato Fossil Beds of Brazil: Window into an Ancient World. Cambridge University Press (Cambridge), pp. 525-533.

Naish, D. 2007. Turtles of the Crato Formation. In Martill, D. M., Bechly, G. & Loveridge, R. F. (eds) The Crato Fossil Beds of Brazil: Window into an Ancient World. Cambridge University Press (Cambridge), pp. 452-457.



Conway, J., Kosemen, C. M. & Naish, D. 2013. Cryptozoologicon Volume I. Irregular Books.

Conway, J., Kosemen, C. M. & Naish, D. 2012. All Yesterdays: Unique and Speculative Views of Dinosaurs and Other Prehistoric Animals. Irregular Books.

Naish, D. 2011. Dinosaur Record Breakers. Carlton Books, London.

Naish, D. 2010. Tetrapod Zoology Book One. CFZ Press, Bideford.

Moody, R. T. J., Buffetaut, E., Naish, D. & Martill, D. M. 2010. Dinosaurs and Other Extinct Saurians: A Historical Perspective. Geological Society, London, Special Publications.

Naish, D. 2010. Dinosaurs Life Size. Barron’s Educational Series, New York.

Naish, D. 2009. The Great Dinosaur Discoveries. A & C Black, London.

Martill, D. M. & Naish, D. 2001. Dinosaurs of the Isle of Wight. The Palaeontological Association, London.

Martill, D. M. & Naish, D. 2000. Walking With Dinosaurs: The Evidence. BBC Worldwide, London.


Book reviews

Naish, D. 2014. [Review of] Pterosaurs, by Mark P. Witton. Princeton University Press, Princeton and Oxford, 2013, 291 pp., ISBN 978-0-691-15061-1. Historical Biology: An International Journal of Paleobiology, DOI: 10.1080/08912963.2014.882099 [pdf here]

Naish, D. 2014. A Review of ‘The Dodo and the Solitaire: A Natural History’. Journal of Vertebrate Paleontology 34, 489-490. [pdf here]

Naish, D. 2012. [Review of] New perspectives on horned dinosaurs: the Royal Tyrrell Museum Ceratopsian Symposium. Historical Biology doi: 10.1080/08912963.2012.688589 [pdf here]

Naish, D. 2012. [Review of] Biology of the Sauropod Dinosaurs: Understanding the Life of Giants edited by Nicole Klein, Kristian Remes, Carole T. Gee, and P. Martin Sander. The Quarterly Review of Biology 87, 53.

Naish, D. 2012. [Review of] Dinosaurs: a field guide/The Princeton field guide to dinosaurs. Historical Biology DOI:10.1080/08912963.2012.670534 [pdf here]

Naish, D. 2011. [Review of] Barnum Brown: the man who discovered Tyrannosaurus rex. Historical Biology DOI:10.1080/08912963.2011.630260 [pdf here]

Naish, D. 2011. [Review of] The second Jurassic dinosaur rush: museums and paleontology in America at the turn of the twentieth century. Historical Biology  DOI:10.1080/08912963.2011.614404 [pdf here]

Naish, D. 2011. [Review of] Glorified dinosaurs: the origin and evolution of birds. Historical Biology 23, 435-438.

Naish, D. 2011. [Review of] The inner bird: anatomy and evolution. Historical Biology 23, 313-316. [pdf here]

Naish, D. 2008. [Review of] Patagonian Mesozoic Reptiles. The Palaeontology Newsletter 62, 92-97.

Naish, D. 2006. The Carnivorous Dinosaurs [review]. The Palaeontology Newsletter 62, 122-126.

Naish, D. 2004. [Review of] African Dinosaurs Unearthed: the Tendaguru Expeditions. Proceedings of the Geologists’ Association 115, 379-380.

Naish, D. 2003. [Review of] The Interrelationships and Evolution of Basal Theropod Dinosaurs. Geological Magazine 140, 729.

Popular and semi-popular articles

Naish, D. 2013. How dinosaurs conquered the world. BBC Focus (Feb’ 2013) 251, 54-59.

Naish, D. 2012. Questions at the frontiers of… dinosaur science. BBC Focus October 2012, 32-33.

Naish, D. 2012. Palaeontology bites back… (response to Ford’s article on alleged aquatic habits of dinosaurs) Laboratory News May 2012, 31-32. [pdf here]

Naish, D. 2012. Should we give up looking for Bigfoot? BBC Focus March 2012, 27.

Naish, D. 2010. Will it float? Scientific American 304 (1), 22. [pdf here]

Naish, D. 2008. Intelligent dinosaurs. Fortean Times 239, 52-53.

Naish, D. 2005. Fossils explained 51: sloths. Geology Today 21 (6), 232-238. [pdf here]

Naish, D. 2004. Fossils explained 48. Placodonts. Geology Today 20 (4), 153-158. [pdf here]

Naish, D. 2004. Fossils explained 46. Ancient toothed whales. Geology Today 20 (2), 72-77. [pdf here]

Naish, D. 2002. Thecocoelurians, calamosaurs and Europe’s largest sauropod: the latest on the Isle of Wight’s dinosaurs. Dino Press 7, 85-95.

Naish, D., Noe, L. F. & Martill, D. M. 2001. Giant pliosaurs and the mysterious ‘Megapleurodon’. Dino Press 4, 98-103.

Naish, D. 2001. Eotyrannus lengi, a new coelurosaur from the Isle of Wight. Dino Press 5, 82-91.

Naish, D. 2001. Fossils explained 34: Crocodilians. Geology Today 17 (2), 71-77. [pdf here]

Naish, D. 2000. 130 years of tree-climbing dinosaurs: Archaeopteryx, ‘arbrosaurs’ and the origin of avian flight. The Quarterly Journal of the Dinosaur Society 4 (1), 20-23. [pdf here]

Naish, D. 2000. Where be monsters? (sea serpents). Fortean Times 132, 40-44.

Naish, D. 1999. Fossil Explained 27: Theropod dinosaurs. Geology Today 15, 234-239. [pdf here]

Naish, D. 1998. Birds of a feather. Fortean Times 108, 34-37.

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